Kundrát, M., Cruickshank, A.R.I., Manning, T.W., and Nudds, J. 2008. Embryos of therizinosauroid theropods from the Upper Cretaceous of China: diagnosis and analysis of ossification patterns. Acta Zoologica 88
ABSTRACT: Exceptionally complete, in ovo dinosaur embryos from the Upper Cretaceous of China are analysed. Ossification patterns of these embryos suggest that they died during the final third of their development. The therizinosauroid identity of the embryos follows from: (1) an edentulous premaxilla with a sharp downturned edge; (2) dentary with a lateral shelf; (3) teeth with fan-shaped crowns, with a few marginal cusps; (4) humerus with a massive deltopectoral crest extending proximally, with a pointed proximomedial tuberosity; (5) ilium with an expanded and hooked preacetabular process; (6) strongly curved hypertrophied manual unguals tapering to sharp points. These embryos are closest to two Chinese therizinosauroids, Neimongosaurus yangi Zhang et al. 2001 and Erliansaurus bellamanus Xu et al. 2002. An elongated narial opening, reduced basipterygoid process, low cervical neural spines, a transversely narrow pubic apron, and a pubic foot expanded anteriorly are found in these embryos and are synapomorphies uniting the Therizinosauroidea and the Oviraptorosauria. Fusion of cervical and caudal neural arches and centra, complete ossification of thoracic ribs and ilium, possible co-ossification of tibia and fibula, fused pubes, complete meta- and acropodial elements, together with small portions of unossified epiphyses of long bones suggest an advanced precociality of these embryos.
Vouz avez remarqué que cette formation la a produit des grosses betes? Achillobator, Electrosaurus, Erliansaurus et encore cet gros oviraptor, pas vraiment l'endroit ideal pour Josephine l'ange gardien se promener
The pectoral girdle and forelimb of the primitive therizinosauroid Falcarius utahensis (Theropoda, Maniraptora): analyzing evolutionary trends within Therizinosauroidea LINDSAY E. ZANNO, Journal of Vertebrate Paleontology, 2006, 26(3):636–650
The recent discovery of a dense, paucispecific bonebed from the Early Cretaceous Cedar Mountain Formation, central Utah, has yielded new information on the morphology and evolution of therizinosaurs. Detailed description of the pectoral girdle and forelimb of Falcarius utahensis—the predominant taxon recovered from this site— provides the basis for a species-level, phylogenetic investigation of this enigmatic group. The analysis, consisting of 32 characters arrayed among 13 taxa, supports previous assertions that Falcarius represents the basal-most known member of the clade. The analysis further supports a monophyletic Therizinosauroidea on the basis of seven unambiguous synapomorphies of the pectoral girdle and forelimb. Contrary to previous hypotheses, analysis of the pectoral girdle and forelimb suggests that Therizinosauridae is more appropriately defined as a derived clade including Nothronychus,Erlicosaurus, Neimongosaurus, Therizinosaurus, and Segnosaurus. Equally strong support is recovered for a clade containing these five genera plus “Alectrosaurus” and Erliansaurus. The morphology of primitive therizinosauroids—characterized by Falcarius, Beipiaosaurus, and Alxasaurus—suggests that Early Cretaceous taxa already exhibited the beginnings of a trend toward increased robustness and altered range of motion of the pectoral girdle and forelimb. Derived therizinosaurs exhibit an amplification of these evolutionary trends as well as increased dorsal reach, increased wrist flexibility, and severe reduction in manual digit length. The functional reorganization of the pectoral girdle and forelimb throughout the evolutionary history of therizinosaurs can be reasonably attributed to a shift from obligatory predation to a novel paleoecological role that reached its pinnacle in derived Late Cretaceous members.
http://download.yousendit.com/D0DF53F037E744CC
Revision of the Tendaguru sauropod dinosaur Tornieria africana (Fraas) and its relevance for sauropod paleobiogeography
KRISTIAN REMES, Journal of Vertebrate Paleontology, 2006, 26(3):651–669
The incompletely known sauropod Tornieria africana from the Upper Jurassic (Tithonian) of Tendaguru, Tanzania, has for over 80 years been regarded to represent a Gondwanan species of the North American genus Barosaurus Marsh (Morrison Formation: Kimmeridgian-Tithonian), but this identification has recently been questioned. The holotype and referred specimens are redescribed here, and the characters present are reevaluated in light of current knowledge of sauropod phylogeny. Synapomorphies of the skull (prefrontal with triangular posterior process) and anterior caudal vertebrae (procoelous centra, presence of diapophyseal laminae, and presence of a pleurocoel) indicate that the Tendaguru material represents a member of the Diplodocinae (Diplodocoidea, Diplodocidae) and is therefore very closely related to Barosaurus and Diplodocus. It differs from all other diplodocine genera in several characters, such as robust anterior caudals with pleurocoels located in the upper third of the centrum and ventral excavations, and stout hind limb proportions similar to Apatosaurus (tibia:femur length ratio less than 0.64). In a phylogenetic analysis, the African form consistently emerges as the sister taxon to Barosaurus+Diplodocus. Therefore, previous suggestions that Tornieria africana is the available name for this taxon are supported by this analysis. The existence of this form in Gondwana contradicts the idea of Laurentian endemism of the diplodocid clade, and is best explained by a vicariance model of diplodocoid paleobiogeography. This implies an extensive ghost lineage of this group, extending back at least as far as the upper Middle Jurassic.
- Mongolie:
# Omnogov:
a) Bayanshiree - CEN/SAN = Erlikosaurus andrewsi, Segnosaurus galbinensis
b) Nemegt - MAA = Therizinosaurus cheloniformis
c) White Beds - MAA = Therizinosaurus cheloniformis
Soudan:
# Province du Nord:
a) Wadi Milk formation (ALB/CEN): Dromaeosauridae indet.
IV- Europe: Angleterre:
# Wight:
a) Wessex formation (BAR): Ornithodesmus cluniculus, Dromaeosauridae indet., Velociraptorinae non nommé
Danemark:
# Bornholm Amt:
a) Jyddegaard formation (VAL/BAR): Dromaeosauroides bornholmensis
Espagne:
# La Rioja:
a) Castellar formation (HAU/BAR): Dromaeosauridae indet.
b) Camarillas formation (BAR): Dromaeosauridae indet.
c) Artoles formation (BAR): Dromaeosauridae indet.
# Cuenca:
a) Calizas de la Huergina formation (BAR): Dromaeosauridae indet ?
# Burgos:
a) Unit S3U1 (CAM): Dromaeosauridae indet.
b) Calizas de Lychnus formation (MAA): Dromaeosaurus indet.
# Huesca:
a) Tremp formation (MAA): Dromaeosauridae indet.
France:
# Aude:
a) Grès de saint-Chinian (CAM/MAA): Variraptor mechinorum
b) Marnes rouges inférieures (MAA): Dromaeosauridae indet.
c) Marnes rouges de Roquelongue (MAA): Dromaeosauridae indet.
# Bouches-du-Rhône:
a) Grès à reptiles (MAA): Variraptor mechinorum, Pyroraptor olympius
# Gard:
a) formation sans nom (MAA): Dromaeosauridae indet ?
b) formation sans nom (CS): Dromaeosauridae indet ?
# Haute-Garonne:
a) Lestaillats Marls (MAA): Dromaeosauridae indet ?
# Hérault:
a) formation sans nom (CAM): Dromaeosauridae indet.
b) Grès de Saint-Chinian (CAM/MAA): Variraptor mechinorum
# Var:
a) Grès à reptiles (MAA): Variraptor mechinorum
Portugal:
# Coimbra:
a) formation sans nom (CAM/MAA): Euronychodon portucalensis
# Leiria:
a) Camadas de Guimarota (KIM): Dromaeosauridae indet.
Roumanie:
# Hunedoara:
a) Sanpetru formation (MAA): Dromaeosauridae indet.
b) Densus Ciula formation (MAA): Dromaeosauridae indet.
abrosaurus: r.délicat
agnosphitys :aux origines inconnues
aletopelta : armure errante
altirhinus au nez haut
amurosaurus
anasazisaurus
animantarx forteresse vivante
bagaraatan petit prédateur
bambiraptor :bambi voleur
beelemodon
bellusaurus beau reptile
besanosaurus
bradycneme jambe lourde
bugenasaura R. à large joue
caenagnathasia mâchoire récente d'Asie
cardiodon dent en forme de coeur
chuanjiesaurus R. de Chuanjie
claorhynchus bec cassé
crichtonsaurus R. de Crichton
cristatusaurus R.crêté
deltadromeus : coureur du Delta
dilong dragon empereur
dinhelrosaurus r.de Dinheiro
domeykosaurus r.de Domeyko
draconyx griffe de Dragon
eolambia avant les Lambeosaurinés
equijubus (de la Montagne de) La Crinière du Cheval
erliansaurus r. d'Erlian
falcarius
gargantuavis
gastonia Pour Robert Gaston
gojirasaurus r. godzilla
gwyneddosaurus